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Special:Whatlinkshere produces some invalid HTML markup. For an example see

http://en.wikipedia.org/w/index.php?title=Special:WhatLinksHere&target=Amazon_River

Now the syntax is:

<ul>
  <ul>
    <li></li>
   </ul>
</ul>

it should be:

<ul>
  <li>
    <ul>
      <li></li>
     </ul>
  </li>
</ul>

I think the error occured in r33602

fixed in r34078.